Bioenergetics, Photosynthesis: Role of CO2 & H2O, Niel's Hypothesis (Lecture 6 Ch 4) NBF Biology 11

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[@BiologyGuardian]

This role of water in photosynthesis was first reported by Van Niel in 1930. He hypothesized that plants split water as a source of hydrogen, releasing oxygen as a byproduct. This observation was based on investigations of photosynthesis in bacteria that make carbohydrates from carbon dioxide, but do not release oxygen. Neil's hypothesis was confirmed in 1940, when for the first time 18O in biological research was used. In first experiment water was made of 18O. The water tagged 18O was added to an alga suspension. The oxygen, evolved during photosynthesis, was found to be radioactive. It was separated and identified. In another experiment carbon dioxide with tagged 18O was added. The oxygen evolved contained none of the isotopes. Thus the source of evolved oxygen was proved to be water. In the following summary, red denotes labelled atoms of Oxygen 18O.

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